Sometime between 3 million and 1.8 million years ago, a part of our genetic population branched off from us and preceded the rest of Early Homo out of Africa into the wide world. The proof of this is in certain 3.1 million year old introgressed genes found in South Asia and the Pacific today, in such fossils as the Hobbit and Meganthropus SE Asia. When homo sapiens first entered parts of Eurasia and SE Asia, they found a branch of hominids already living there who'd left our direct lineage shortly after we'd enjoyed our last tryst with the proto-chimpanzees. This first wave of Early Homo entered South Asia two million years ago or more, before Homo ergaster and Homo erectus had even evolved. Some of their descendants lived in isolation like the Indonesian hobbits, and survived into the late Paleolithic, if not longer. Others have been assimilated into wave after wave of other hominids over the past 2 million years, the majority of their genes having been selected against.
The Madrasian tools of India provide further evidence of these peoples' movements out of Africa and across Eurasia. This lithic culture is unmistakably Mode I, which is only known to have been used by Early Homo and Australopithecines. But it begins over 1.5 million years ago in South Asia and continues until 10,000 years ago in isolated areas.
Though no fossil evidence remains of the Madrasian tools' makers, these hominids would have shared traits and brain size with Homo Habilis. Some variations of Eurasian members of Early Homo include Homo Georgicus, Homo erectus modjokertensis (Taung Child), and Meganthropus Robustus. Several more candidates have recently been found in East Asia and the Phillipines.
Our ancestors had no particular advantage over these hominids when they first left Africa. But sometime around 2.2 million years ago our clan developed a new brain gene that gave us a little bit of an edge over everybody else, so we started expanding faster than everyone else,and incorporating everyone else into our population and culture while simultaneously outbreeding them. The first evidence we find of this expansion is Homo Ergaster, who appears with a more advanced type of tool in Eastern Africa around 1.8 million years ago.
Researchers call part of this expansion the Acheulian, but it extended far beyond the area where that particular tool tradition is found.
The early hominids who had proceeded us out of Africa were mostly assimilated in the wave of this expansion, but some of them managed to avoid the Acheulian expansion and lived separately from our direct ancestors in South Asia and SE Asia until the late paleolithic...and possibly even into historic times. We will call these the Hobbit in South-East Asia and Homo Vanara in South Asia, after the Vedic word for the forest dwelling ape-men of southern India.
Shortly after the appearance of Homo Ergaster in Africa, Acheulian tools appeared in India. However, the older tool traditions persist in South Asia alongside these newer ones. Unfortunately, there are no fossils of either tool-making species from this time from South Asia.
The fossil of Homo erectus modjokertensis shows up on Java around 1.8 million years ago, slightly earlier than the dating of the first Madrasian tools. But the Mode I lithic industry there is akin to neither of its Western counterparts. This might be due to a difference in resources, as SE Asia has less flint and more chert and bamboo. The modjokertensis hominid was a child when it died, and therefore not fully developed, but its morphology seems intermediary between Early Homo and the later Homo Erectus
Fossils of the sister species of Homo Ergaster, Homo Erectus, appear in South East Asia around 1.49 million years ago . While some of the Javan fossils seem even more advanced than homo ergaster, others from this time in SE Asia harken back to Modjokertensis, Georgicus, Habilis, and even the Australopithecines. Meanwhile, specimens of this age from Africa are rarely referred to as ergaster anymore, but take on the erectus moniker as well. Sometimes the catch-all term used is Homo eregasterectus.
But from 1.4 to 1 million years ago, Africa looks to have been all but abandoned, at least for intermittent periods that probably coincide with temperature cycles. There are no fossils and few tools to mark the presence of eregasterectus, and this may coincide with hot periods during an interglacial when the Sahara expanded to make at least the northern and eastern part of the continent inhospitable. However, we know that Africa was not completely devoid of hominins at this time, because genetic evidence shows that between 1.3 and 1.2 million years ago, a population of Homo ergasterectus separated itself from our gene pool. They remained in isolation somewhere in Africa until being assimilated by the Hadza pygmies (or their immediate ancestors) over a million years later. We know this because the Hadza tribes alone possess these 1.3 million year old gene variants, and studies show they entered the Hadza population roughly 50,000 to 100,000 years ago.
Around the same time this African lineage broke off from our main lineage and isolated itself somewhere in Africa, hominid bones appear in Turkey and Spain for the first time. They are more akin to early Ergaster and Georgicus than to Homo ergasterectus, and are associated with Mode I Oldowan tools This suggests to many that they were not part of the spreading culture that used Mode 2 or Acheulian tools. Rather, they represent members of Early Homo who didn't get the Mode 2 memo before heading into Europe out of Africa.
The fossils on Java become more plentiful and diverse during this proposed "hot period," including the first finds of Java Man. This subspecies is perhaps more closely related to Homo Ergasterectus than any other hominid of 1.4 million years ago, and is the type specimen for "Classic Erectus." However, Java man shared his island with other more basal forms, such as Meganthropus, which might share affinities with Homo Habils and even the Austrolipithecines. Other less robust specimens that resemble the earlier Modjokertensis, Georgicus, and Habilis, also lived on Java during this period, and possibly continued living there until 50,000 years ago or later.
It could be that what we are seeing on Java at this time is a snapshot of what was happening in the whole of Sundaland just before 1.4 million years ago, when the continent had been above sea level. The whole of Java would have been undesirable highlands until the sea rose, potentially stranding several species of SE Asian hominid together.
In any event. around 1.1 million years ago yet another population separated itself from our direct ancestral genomic population. This was the Microcephalin D hominid, who we will call "Classic Erectus," and it did not recombine with our own genome until around 37,000 years ago. Classic Erectus could also be responsible for some of the introgressed genes of the "Mystery Hominid" present in Denisovans, Malanesians, SE Asians, and some South Asians. This population must have had at least some genetic exchange with the Hobbit or Homo Vanara, since "Mystery Hominid" introgression into the aforementioned populations often comes with genes from the 3 million year old divergence of Homo.
It can be seen as a far shore of the genomic wave in which Homo Ergasterectus brought SRGAP2C to the Early Hominids spread across Eurasia who only had SRGAP2B. The Los Huesos, who's mtDNA is 450,000 years older that of the "Denisovan genome," may also be evidence of this sweep, as are the Denisovans themselves. Classic Erectus broke off about 1.1 million years ago, the mitochondrial ancestor of Los Huesos broke of around 1.05 million years ago, and the oldest (possibly introgressed) mtDNA haplogroups of Denisovan broke off about a million years ago.
However, some Early Homo populations must have avoided the SRGAP2C sweep, because we have 3 million year old introgression without the accompanying "Mystery Hominid" or "Denisovan" in a few people living in Western South Asia today. Plus, we have the Hobbits of South East Asia who's morphology suggests an initial split from our ancestral population at around the same time.
The distribution of microcephalin D in modern humans suggests that the hominid that originally carried it lived somewhere between Papua New Guinea and Pakistan, since populations these two areas have the highest amount and its distribution across the globe points to an origin point in either Eastern South Asia or South East Asia.
Could it be that a population of Classic Erectus was stranded on Java or another Sunda island during a hot period of high seas roughly 1.1 million years ago, along with with a population of Hobbits?
If so, some Hobbits apparently escaped across the Wallace Line to Flores right before the hot period, or remained at least somewhat separate from Classic Erectus on another Sunda Island near Java until 780,000 years ago at the beginning of a glacial period when seas would have dropped again and fossils of the Hobbit's ancestors actuall start appearing on Flores.
So at this point, we have two species of Early Homo and one species of Classic Erectus living in South Asia and/or South East Asia. We have our genome of Homo Ergasterectus living Betwee or in South Africa and India. We have a species of Homo Ergasterectus living in African isolation, and we have a species of early Ergasterectus in Europe as well.
1 million years in the date of the diversion of the oldest of the Denisovan mtdna haplogroups, and these are only four hundred and fifty thousand years younger than the mtdna of Los Huesos. The nuclear DNA of Denisovan is branched off from us hundreds of thousands of years later. What this tells us is that Los Huesos females (and probably Classic Erectus females) were part of a genetic sweep that replaced Early Homo mtDNA haplogroups across Eurasia, and Denisovan females are the result of a later part of that same sweep that replaced Early Homo in China. About 1.1 million years ago, a population branched off from our own drove all Early Homo mtDNA into extinction except for a few small population living in splendid isolation somewhere in S. or SE Asia. Another way to say this is that Denisovan women conquered China before the rest of Denisova did, but in actuality they were probably not true Denisovans until the defining nuclear DNA arrived.
At the same time that the earliest of denisovan mothers we're branching off from our own genomic population, a new brain mutation called SRGAP2D was born. It is probable that the mutation happened when one of our ancestors that were living somewhere in or between africa and South Asia at the time, but we cannot rule out the possibility that arose in one of the aforementioned subspecies of hominid. The gene is present in all modern humans, so if it enter breast before or round 350000 years ago we would not be able to tell whether it introgressed or not. Help the Common Man your modern humans neanderthals and nuclear denisovans where the next to me a year worldwide genomic sweep, we will assume that the former.
Heidelberg Man is the name most often used to describe this common ancestor. The Heidelberg man may have his roots in the SRGAP2D mutation that appeared 1 million years ago, fossils don't show up until around 600,000 years ago,
As expected, they show up on the outer borders of the area I've proposed for our own ancestors: the Narmada Valley in North West India and East Africa. Narmada Man and the Bodo and Kabwe craniums have a gigger brain size than any hominid that has come before them, and share traits with later "Classic Heidelbergs" from Africa and Eurasia. However, Homo Ergasterectus material is also appearing in east Africa at this time, having begun reappearing in the fossil record around 1 million years ago after a 400,000 year hiatus.
Only slightly later, the cranium of Ceprano Man is deposited in Europe. It is highly divergent from the antecessor fossils that precede it there and has more affinities with Classic erectus and its contemporary Narmada Man than it has with Kabwe and Bodo, but shares a resemblance to later Classic Heidelberg.
This bears all the markings of a diffusion of an advanced hominid from South West Asia or the Middle East, who we will call Classic Heidelberg.
Speaking of the ancestral population that loud between africa europe central Asia and India drum a million years 2 sometime after 200000 years ago, I do not see that they look much like classic heidelberg. They were likely comparatively short, with a head shape more like classic erectus. Mixing wheel homo ergaster and Ergasterectus to the immediate Wells and North is probably what is classic heidelberg the tall stature, more rounded cranium, and neonatal face as it radiated into Africa and Europe. It may also heidelberg interred europe from northeast, germany and the British Isles had not yet made it with antecessor, goodwill in southern Europe previous and seems to be the word neanderthal. Perhaps even a separate group of early Heidelberg, Ceprano Man Man, entered from the southeast, and the three groups plus the hardships of a later Ice Age is what begat Classic Neanderthal in Europe. Nneka call me many signs of hybridization between 600000 and 250000 years ago in the fossil record, until Classic Neanderthal appears and becomes ubiquitous there.
The greatest evidence we have for them entergy the Los huesos specimen. It has mtdna from the earliest wave of denisovan expansion, but it's nuclear DNA is decidedly Neanderthal. Heidelberg entered Europe around 600,000 years ago and assimilated Antecessor, a sister species of "early Denisovan," and drove the earlier mtDNA haplogroups there to extinction.
Neanderthal/ Denisovan mtdna split off from our own about 840000 years ago, and then split into Neanderthal and Denisovan respectively about 640,000 years ago, suggesting a period of isolation of 200 thousand years, probably somewhere in Central Asia, which is inhabited intermittently during this period depending on glaciation and climate. It is also likely that a population from this group made its way into Pakistan and India during this time, or was already there to begin with, judging from the similarity of Neanderthal and western S. Asian lithics and the current distribution of neanderthal genes in modern humans.
Rather than saying that neanderthals and the larger part of denisovans broke away from Hss 840,000 years ago, it may be more useful to say that we broke away from them it, and lived in an area that encompassed North and East Africa and South West Asia as our siblings spread out across the world.
700000 years ago yet another population Branch out from our own, heading West and South into greater Africa. It is likely that this subspecies of Heidelberg assimilated the aforementioned African archaic that had been living there since 1.3 million years ago, as introgression from the group in pygmies often contains these older genes as well.
600,000 years ago another group split off from Heidelberg, heading East. Their genes are now only found at low levels in the Andaman islanders. The appearance of "pygmy heidelbergs" in northern India around 150,000 years ago may herald their arrival.
400000 years ago one group of Heidelberg left our gene pool and migrated into Subsaharan or West Africa and is represented by the later introgressed Y Haplogroup A00, which is today only found in a handful of people with ancestry from that region. It is separated by 200 thousand years from the haplogroups of the rest of modern man, suggesting 200 thousand years of haplogroup extinctions from which only two haplogroups among millions survived. Shortly after the divergence of this haplogroup, the tallest examples of Classic Heidelberg appear in Africa in the form of Rhodesian Man and Homo Sapiens Idaltu. Some populations are thought to have averaged 7 foot among males. African Ergaster happens to be the tallest member of the Genus Homo before African Heidelbergs such as Rhodesian man and Idaltu (all are 6'1+), while non-African Heidelbergs are the same height as the erectus that went before them (Antecessor and Los huesos are 6', all other non-African Erectus and "Heidelberg" range between 3' and 5'10, with 5'10 being an extreme rather than a population average). In evolutionary thought, it is considered easier to shrink than to grow, although I would guess that environment, gene exchange, and circumstance might also have something to do with it.
The last divergence from our from own direct ancestral population (that we know about) occurred between 250 and 350 thousand years ago and is evidenced in the mtDNA of an anatomically modern human dated 40-60 thousand years ago in Australia known as Mungo Man.
Anatomically humans show up in the fossil record until 160,000 years ago, and they are all in Africa, which might seem to contradict the DNA. Research agree that though they were both homo Sapien sapiens, neither our Mito eve nor that of Mungo Man were likely anatomically modern. Some hold that anatomically modern Homo Sapien Sapiens don't actually appear until around 120,000 years ago or later. How can it be that two seperate gene pools that were not anatomically modern to begin with both become anatomically modern by 45000 years ago? For that matter, since Y Haplogroup A00 split off from our own 400,000 years ago, when Homo Sapien Sapiens wasn't even around, how can A00 people living today be anatomically modern?
The answer is interbreding between the groups. According to john Hawks and the bBC program first Peoples, Anatomically modern humans formed from genetic interchange between several different groups of heidelbergensis. We know this because different groups of Archaic Humans living in or near Africa from 350 thousand to 90 thousand years ago have different pieces of the puzzle that coalesced to creat anatomically modern human morphology. The group that included the artist formerly known as Y DNA Adam, the group that contained Y Haplogroup A00, the group that contained Mitochondrial Eve, and the group that contained LM3 eve were all subgroups of Heidelberg, with different traits, and they came together to form the traits inherent in every living person today. Y Adam and Mito Eve just won the lineage marker part of the genome, there is no reason to believe that other groups didn't win other parts. Any introgression from the LM3 or A00 populations that happened before anatomically modern human came to be would not be seen as introgression, because everybody on Earth has them today. Too, unlike introgression from neanderthals and Denisovans, nuclear material from the A00 and LM3 populations would only be separated from our own nuclear material by a couple of hundred thousand years or less, and equidistant from that of Neanderthals and Denisovans, so there is even the possibility of introgression after the time in which they coelesced into AMH. We even have evidence for that in the form of a erroneous insert into 39% of modern human's own chromosome 11 that looks just like mtDNA from LM3's Eve. If not for this insert, scientists would not know that our own ancestors even bred with our Pacific sister species, leaving the possibility that there were still other extinct populations of Homo sapien sapiens and AMH that bred with us but remain undetected.
It is even a possibility that all of our Archaic Introgression stems from these "Ghost Population" having assimilated neanderthals, denisovans, and others before we assimilated them in turn.
The extreme of this is that all members of the Genus Homo should be considered Homo sapien sapiens, but because there was likely limited fertility between some of these groups and because there is cause for separation on anatomical grounds, not to mention the unlikelyhood that a modern human could successfully mate with a cloned Homo Habilis, I tend to disagree with this preference. However the various species and subspecies of the genus homo world without a doubt a part of an extended gene pool promise augmented and recombined many times over the past 3.5 million years.
90,000 years ago, LM3 may even still been a part of our own gene pool, and A00's population may not have yet recombined with it yet.
That's when a cold spell started pushing neanderthals into the Middle East and split our population into at least two parts. LM3 and possibly several Y and mtDNA haplogroups that survived into the present (Y Hap C and mtDNA M?) survived in South China where there is presence is known from 120,000 years ago, while the descendents of mito Eve (us) were limited to Africa. In a territory from Africa to Pakistan, and maybe into India, Neanderthal ruled and assimilated the AMH and Heidelberg remained in the region for the next 50 thousand years. we have evidence of this in not only the fossil record but from the introgressed genes of homo sapien sapiens found in the fossils of this self-same group of neanderthals, and their introgression into us.
This same cold spell may have opened up the land bridge to the Andaman Islands for the Early Heidelberg that is Andamanese Archaic Introgression. This subgroup of Heidelberg branched off from our own ancestors 600,000 years ago. By 150,000 years ago, a "pygmy" form of Heidelberg shows up in Northern India and lives for the next 30 thousand years or more alongside a taller but more gracile subgroup that had lived there for at least 600,000 years before. Their size and certain anatomical traits are almost identical to Andaman Islanders. Andaman islander Introgression may even be a combination of these two people's genes, with the 600,000 year old fossils, tools and introgressed genes being the taller Heidelbergs and the 150,000 year old tools and fossils being the Neanderthal Introgression in Andaman Islanders. It may also be that this group included women of the LM3 haplogroup, or a ghost population of Hss that carried the LM3 insert, because Andaman Islanders have it too. It could even have included Y Haplogroup C and D and mtDNA M, but this is doubtful, and here is the reason: introgression is only a fraction of Andaman Islander DNA. The vast majority of it is from the Homo Sapien Sapiens genome, and their maternal and paternal haplogroups are from within the even more finite branches of Y haplogroup A and mtDNA Eve. More importantly, they have all of the "updates" that make us human, unlike Indian Pygmy Heidelberg/Neanderthal or any Hominid that branched off from us 600,000 years ago would have. The dates just don't compute- C,D, and M are only around 60 thousand years old. their age would have to be multiplied by three to comply with the dates of the first pygmy Heidelberg's arrival in Northern India! Furthermore, the Andaman islanders are thought to have obtained their 600,000 year old genes only 35,000 years ago, and their Neanderthal genes not long before....and certainly not over 125,000 years before.
But what if the original population from which Andaman Islanders received their introgression was part Hss in the first place? What if in addition to neanderthal and 600,000 year old "Heidelberg genes" it also contained an equal or greater portion of genes from LM3 or Mito Eve? What if it contained Y Haplogroup A0? What if the shared cultural and physical traits of South East Asian Negritos and African pigmy's, such as steatopygia and the similarities between the two groups in cranial morphology as reported in the last study on the subject conducted in 1973.
What if the biggest difference between the negritos and pygmies and ourselves isn't the amount of neanderthal Introgression or admixture from 600,000 and 700,000 year old early heidelbergs. Maybe it's that they were never fully assimilated by the taller group of Homo Sapien sapiens who that once shared an enviroment with in the original homeland of Homo Sapien Sapiens, two populations enjoying two different niches and occasionally exchanging genes, until the Neanderthals drove both of us out.
It may even be that Y haps C and D and mtDNA M in Andaman Islanders comes from a handful of shipwrecked sailors who entered the gene pool sometime between 35,000 and 10,000 years ago.
More likely, however, is that they a group of Heidelberg/Neanderthal/Hss pygmies lived in Northern India, Southern China, and Indochina from 150,000 years ago to at least 45,000 years ago and shared the landscape and sometimes exchanged genes with a taller hybrid of heidelberg/ Hss hybrid that had experienced less introgression from Neanderthals. I say this because the taller specimens linked to Hss from this time and era are more gracile, and because the probable descendant Mungo Man is hyper-gracile and has supposedly less neanderthal introgression than other Hss specimens outside of Africa from a similar date.
An interesting sidenote is that Hss from both Japan and Australia during this period exhibit the practice of tooth knocking. The Japanese Hss have large denisovan like teeth and are Hyper-Robust like Denisovans and they are short and robust like the Indian Heidelberg pygmies, but the Australians are tall and lean. I suspect that the practice may have resulted from a problem with tooth development during adolescence in hybrids of the two groups, or in hybrids of the two groups and neanderthals or Asian hominids. The practice continued in Japan until the Neolithic Jomon Period, in Australia into the present, and back-migrated into Africa about 20,000 years ago within the late Aterian or Ibero-Maurusian cultures, possibly with Y Haplogroup DE and MtDNA M.
So it looks like we had two groups of Homo sapiens sapiens living in an area africa to indochina from around 150,000 years ago to 90,000 years ago. One exploded plains, savannas, and grasslands and was tall and gracile. The other exploited forests and was short and stocky. Ice sheets and Neanderthals kept them out of Europe and Central Asia and something kept them out of most of India, likely something closely related to neanderthals judging my the Mousterian tools found there. Something was also keeping both groups of Hss out of Sub-Saharan and West Africa, where A00 and Archaic African Introgression were waiting to be assimilated by people living there today. Indeed, archaic fossils persists in west Africa until 10,000 years ago.
A cold spell drove the Neanderthals into SW Asia and the Middle East, and separated the genome of LM3 and Negritos from the genome of Mito Eve and Pygmies. Before 90000 years ago, a few fossils of Homo Sapien sapiens in East Africa, China, and the Levant exhibit chins. After 90000 years ago, chins begin to pop up all over Africa and in Southern Neanderthals too. Some African examples dated between 70 thousand and 90 thousand years ago were considered neanderthals until the 1980s, when they were reassigned the Homo Sapien Sapiens classification due to a number of modern features. It looks for all the world like Neandertha/Hss hybrids from the borders of Africa and SW Asia are beginning an assimilation of the entire continent. The dates match up much well with the back-migration of Y Haplogroup BT, and with a second wave of DE and/or its offspring, basal y haplogroup E. The only Africans who seem to have escaped this assimilation are the pygmies, who likely had higher Malaria immunity due to their archaic introgression, and those who mixed with a population of Heidelbergs that contained A00. Both happened in Subsaharan or West Africa during the Upper Pleolithic, probably around 37,000 years ago.
Today, back-migrated Y haplogroup E makes up over 70% of men native to Africa and the vast majority of Africans outside of Subsaharan Africa possess neanderthal introgression. It is mainly only pygmies who have managed to avoid it, and they have introgression that likely serves them far better than a cold-bred hominid's would.
From 70000 years ago to about 45,000 years ago the Y Hap DE neanderthal/Hss hybrid males dominated most of SW Asia, Northern and Eastern Africa, and the Middle East. Their original mtDNA counterpart was likely N, but they also picked up some mtDNA M, possible from the North Indian Pygmies. Sometime around 20,000 years ago a group of them brought tooth-knocking and mtDNA haplogroup M into Africa, and at some point around this same time one of their groups reached the Andaman Islands carrying the assimilated genes of proto-negritos.
This may have been the result of the population in which de was dominant being split in half, as the genome of homo sapiens sapiens had been split in half by neanderthals before. The reason for this was either the emptying of Central asia and the middle East due to climactic conditions or the incursion of another hybrid group from India, the one that contained Y haplogroup FC.
The dispersal of Y Haplogroup C, probably from South Asia, begins around 60,000 years ago and continues into Eurasia and South east Asia. The dispersal of F begins about 45,000 years ago and does the same. My best guess is that C represents those South Asian Hss that bred with the short woodland Neanderthals of South Asia and that F represents those who bred with the taller neanderthals of South Asia. Hybrid vigor allowed both groups' expansions into larger India, assimilating the early neanderthals there who had already mixed with an incursion Microcephalin D hominids that were expanding in the wake of lower sea levels from South east Asia. This is why the Microcephalin D gene is most prevalent in populations dominated by F haplogroups and mostly absent in African and Asian groups dominated by E and D, the offspring of Y haplogroup DE. Assimilation of the 3 million year divergent Homo Varana probably didn't happen until 21 thousand years ago at the last glacial maximum, when Adam's bridge to Sri Lanka became dry again. The resulting hybrids are known as Balangoda man, and some of them seemed to have moved to SW of India itself where they live to this day.
The push of F and C into central Asia pushed neanderthal East and West, and in the the East they began assimilating their sister species Denisovan, who had already begun assimilating Early Homo some 300 thousand years before. Neanderthal like skulls have been found in Mongolia and near Biejing with neanderthal and other archaic features with dates as late as 9000 B.C. The Red Deer Cave people possess triats from Erectus, neanderthal, and Hss...along with a few that only they can boast. Neanderthal stone balls, previously only found in Europe, show up around 40 thousand years ago in China and continue into South East Asia no later than 27,000 years ago. This is why Northern Chinese, native Amnericans. and Papuans have more Neanderthal DNA than most Europeans: Neanderthal assimilated most of the Denisovan and the Microcephalin D populations before we got there, along with much of the genome of our Mungo Man brother LM3.
It also seems that neanderthals and other hominids liked our females better than their own, as their is evidence for Hss introgression into Neanderthals in some Asian specimens and our mtDNA haplogroups seem to preceed us into certain areas, such as Tibet and Europe, long before any of our surviving Y haplogroups get there. When we took the world back from Neanderthal, we must not have killed the women, or at least we didn't kill the women who looked like us.
But why would all of these dominant neanderthal hybrids who took over the world except for african pygmies always be from an Hss haplogroup rather than a neanderthal one? It may have something to do with mammal pattern infertility. In mammalian hybrids, the females are more likely to be fertile than the males. The lack of neanderthal genital introgression makes it probable that hybrids who inherited the blackneanderthal reproductive genes rather than their Hss parent's were less successful in producing offspring. The other genes missing from the Neanderthal genome in modern introgression have to do with eyesight and may have been a factor in the size of the occipital bun, which could result in birth complications. if Neanderthals were Blood Type O Negative as is supposed, it may be that they could only have one child with an Hss wiothout the aid of modern medicine. It might also be because homo Sapien sapiens was the first hominid that did not need to go into heat in order to procreate. This could explain other subgroups choosing Hss women over their own and the rapid expansion of Hss genes after the Toba catastrophe. We simply outbred them. It could also be that Hss has a more neonatal look than any hominid that preceded it. We were originally pygmies that were just too cute to kill, and our women were smoking hot!
The Madrasian tools of India provide further evidence of these peoples' movements out of Africa and across Eurasia. This lithic culture is unmistakably Mode I, which is only known to have been used by Early Homo and Australopithecines. But it begins over 1.5 million years ago in South Asia and continues until 10,000 years ago in isolated areas.
Though no fossil evidence remains of the Madrasian tools' makers, these hominids would have shared traits and brain size with Homo Habilis. Some variations of Eurasian members of Early Homo include Homo Georgicus, Homo erectus modjokertensis (Taung Child), and Meganthropus Robustus. Several more candidates have recently been found in East Asia and the Phillipines.
Our ancestors had no particular advantage over these hominids when they first left Africa. But sometime around 2.2 million years ago our clan developed a new brain gene that gave us a little bit of an edge over everybody else, so we started expanding faster than everyone else,and incorporating everyone else into our population and culture while simultaneously outbreeding them. The first evidence we find of this expansion is Homo Ergaster, who appears with a more advanced type of tool in Eastern Africa around 1.8 million years ago.
Researchers call part of this expansion the Acheulian, but it extended far beyond the area where that particular tool tradition is found.
The early hominids who had proceeded us out of Africa were mostly assimilated in the wave of this expansion, but some of them managed to avoid the Acheulian expansion and lived separately from our direct ancestors in South Asia and SE Asia until the late paleolithic...and possibly even into historic times. We will call these the Hobbit in South-East Asia and Homo Vanara in South Asia, after the Vedic word for the forest dwelling ape-men of southern India.
Shortly after the appearance of Homo Ergaster in Africa, Acheulian tools appeared in India. However, the older tool traditions persist in South Asia alongside these newer ones. Unfortunately, there are no fossils of either tool-making species from this time from South Asia.
The fossil of Homo erectus modjokertensis shows up on Java around 1.8 million years ago, slightly earlier than the dating of the first Madrasian tools. But the Mode I lithic industry there is akin to neither of its Western counterparts. This might be due to a difference in resources, as SE Asia has less flint and more chert and bamboo. The modjokertensis hominid was a child when it died, and therefore not fully developed, but its morphology seems intermediary between Early Homo and the later Homo Erectus
Fossils of the sister species of Homo Ergaster, Homo Erectus, appear in South East Asia around 1.49 million years ago . While some of the Javan fossils seem even more advanced than homo ergaster, others from this time in SE Asia harken back to Modjokertensis, Georgicus, Habilis, and even the Australopithecines. Meanwhile, specimens of this age from Africa are rarely referred to as ergaster anymore, but take on the erectus moniker as well. Sometimes the catch-all term used is Homo eregasterectus.
But from 1.4 to 1 million years ago, Africa looks to have been all but abandoned, at least for intermittent periods that probably coincide with temperature cycles. There are no fossils and few tools to mark the presence of eregasterectus, and this may coincide with hot periods during an interglacial when the Sahara expanded to make at least the northern and eastern part of the continent inhospitable. However, we know that Africa was not completely devoid of hominins at this time, because genetic evidence shows that between 1.3 and 1.2 million years ago, a population of Homo ergasterectus separated itself from our gene pool. They remained in isolation somewhere in Africa until being assimilated by the Hadza pygmies (or their immediate ancestors) over a million years later. We know this because the Hadza tribes alone possess these 1.3 million year old gene variants, and studies show they entered the Hadza population roughly 50,000 to 100,000 years ago.
Around the same time this African lineage broke off from our main lineage and isolated itself somewhere in Africa, hominid bones appear in Turkey and Spain for the first time. They are more akin to early Ergaster and Georgicus than to Homo ergasterectus, and are associated with Mode I Oldowan tools This suggests to many that they were not part of the spreading culture that used Mode 2 or Acheulian tools. Rather, they represent members of Early Homo who didn't get the Mode 2 memo before heading into Europe out of Africa.
The fossils on Java become more plentiful and diverse during this proposed "hot period," including the first finds of Java Man. This subspecies is perhaps more closely related to Homo Ergasterectus than any other hominid of 1.4 million years ago, and is the type specimen for "Classic Erectus." However, Java man shared his island with other more basal forms, such as Meganthropus, which might share affinities with Homo Habils and even the Austrolipithecines. Other less robust specimens that resemble the earlier Modjokertensis, Georgicus, and Habilis, also lived on Java during this period, and possibly continued living there until 50,000 years ago or later.
It could be that what we are seeing on Java at this time is a snapshot of what was happening in the whole of Sundaland just before 1.4 million years ago, when the continent had been above sea level. The whole of Java would have been undesirable highlands until the sea rose, potentially stranding several species of SE Asian hominid together.
In any event. around 1.1 million years ago yet another population separated itself from our direct ancestral genomic population. This was the Microcephalin D hominid, who we will call "Classic Erectus," and it did not recombine with our own genome until around 37,000 years ago. Classic Erectus could also be responsible for some of the introgressed genes of the "Mystery Hominid" present in Denisovans, Malanesians, SE Asians, and some South Asians. This population must have had at least some genetic exchange with the Hobbit or Homo Vanara, since "Mystery Hominid" introgression into the aforementioned populations often comes with genes from the 3 million year old divergence of Homo.
It can be seen as a far shore of the genomic wave in which Homo Ergasterectus brought SRGAP2C to the Early Hominids spread across Eurasia who only had SRGAP2B. The Los Huesos, who's mtDNA is 450,000 years older that of the "Denisovan genome," may also be evidence of this sweep, as are the Denisovans themselves. Classic Erectus broke off about 1.1 million years ago, the mitochondrial ancestor of Los Huesos broke of around 1.05 million years ago, and the oldest (possibly introgressed) mtDNA haplogroups of Denisovan broke off about a million years ago.
However, some Early Homo populations must have avoided the SRGAP2C sweep, because we have 3 million year old introgression without the accompanying "Mystery Hominid" or "Denisovan" in a few people living in Western South Asia today. Plus, we have the Hobbits of South East Asia who's morphology suggests an initial split from our ancestral population at around the same time.
The distribution of microcephalin D in modern humans suggests that the hominid that originally carried it lived somewhere between Papua New Guinea and Pakistan, since populations these two areas have the highest amount and its distribution across the globe points to an origin point in either Eastern South Asia or South East Asia.
Could it be that a population of Classic Erectus was stranded on Java or another Sunda island during a hot period of high seas roughly 1.1 million years ago, along with with a population of Hobbits?
If so, some Hobbits apparently escaped across the Wallace Line to Flores right before the hot period, or remained at least somewhat separate from Classic Erectus on another Sunda Island near Java until 780,000 years ago at the beginning of a glacial period when seas would have dropped again and fossils of the Hobbit's ancestors actuall start appearing on Flores.
So at this point, we have two species of Early Homo and one species of Classic Erectus living in South Asia and/or South East Asia. We have our genome of Homo Ergasterectus living Betwee or in South Africa and India. We have a species of Homo Ergasterectus living in African isolation, and we have a species of early Ergasterectus in Europe as well.
1 million years in the date of the diversion of the oldest of the Denisovan mtdna haplogroups, and these are only four hundred and fifty thousand years younger than the mtdna of Los Huesos. The nuclear DNA of Denisovan is branched off from us hundreds of thousands of years later. What this tells us is that Los Huesos females (and probably Classic Erectus females) were part of a genetic sweep that replaced Early Homo mtDNA haplogroups across Eurasia, and Denisovan females are the result of a later part of that same sweep that replaced Early Homo in China. About 1.1 million years ago, a population branched off from our own drove all Early Homo mtDNA into extinction except for a few small population living in splendid isolation somewhere in S. or SE Asia. Another way to say this is that Denisovan women conquered China before the rest of Denisova did, but in actuality they were probably not true Denisovans until the defining nuclear DNA arrived.
At the same time that the earliest of denisovan mothers we're branching off from our own genomic population, a new brain mutation called SRGAP2D was born. It is probable that the mutation happened when one of our ancestors that were living somewhere in or between africa and South Asia at the time, but we cannot rule out the possibility that arose in one of the aforementioned subspecies of hominid. The gene is present in all modern humans, so if it enter breast before or round 350000 years ago we would not be able to tell whether it introgressed or not. Help the Common Man your modern humans neanderthals and nuclear denisovans where the next to me a year worldwide genomic sweep, we will assume that the former.
Heidelberg Man is the name most often used to describe this common ancestor. The Heidelberg man may have his roots in the SRGAP2D mutation that appeared 1 million years ago, fossils don't show up until around 600,000 years ago,
As expected, they show up on the outer borders of the area I've proposed for our own ancestors: the Narmada Valley in North West India and East Africa. Narmada Man and the Bodo and Kabwe craniums have a gigger brain size than any hominid that has come before them, and share traits with later "Classic Heidelbergs" from Africa and Eurasia. However, Homo Ergasterectus material is also appearing in east Africa at this time, having begun reappearing in the fossil record around 1 million years ago after a 400,000 year hiatus.
Only slightly later, the cranium of Ceprano Man is deposited in Europe. It is highly divergent from the antecessor fossils that precede it there and has more affinities with Classic erectus and its contemporary Narmada Man than it has with Kabwe and Bodo, but shares a resemblance to later Classic Heidelberg.
This bears all the markings of a diffusion of an advanced hominid from South West Asia or the Middle East, who we will call Classic Heidelberg.
Speaking of the ancestral population that loud between africa europe central Asia and India drum a million years 2 sometime after 200000 years ago, I do not see that they look much like classic heidelberg. They were likely comparatively short, with a head shape more like classic erectus. Mixing wheel homo ergaster and Ergasterectus to the immediate Wells and North is probably what is classic heidelberg the tall stature, more rounded cranium, and neonatal face as it radiated into Africa and Europe. It may also heidelberg interred europe from northeast, germany and the British Isles had not yet made it with antecessor, goodwill in southern Europe previous and seems to be the word neanderthal. Perhaps even a separate group of early Heidelberg, Ceprano Man Man, entered from the southeast, and the three groups plus the hardships of a later Ice Age is what begat Classic Neanderthal in Europe. Nneka call me many signs of hybridization between 600000 and 250000 years ago in the fossil record, until Classic Neanderthal appears and becomes ubiquitous there.
The greatest evidence we have for them entergy the Los huesos specimen. It has mtdna from the earliest wave of denisovan expansion, but it's nuclear DNA is decidedly Neanderthal. Heidelberg entered Europe around 600,000 years ago and assimilated Antecessor, a sister species of "early Denisovan," and drove the earlier mtDNA haplogroups there to extinction.
Neanderthal/ Denisovan mtdna split off from our own about 840000 years ago, and then split into Neanderthal and Denisovan respectively about 640,000 years ago, suggesting a period of isolation of 200 thousand years, probably somewhere in Central Asia, which is inhabited intermittently during this period depending on glaciation and climate. It is also likely that a population from this group made its way into Pakistan and India during this time, or was already there to begin with, judging from the similarity of Neanderthal and western S. Asian lithics and the current distribution of neanderthal genes in modern humans.
Rather than saying that neanderthals and the larger part of denisovans broke away from Hss 840,000 years ago, it may be more useful to say that we broke away from them it, and lived in an area that encompassed North and East Africa and South West Asia as our siblings spread out across the world.
700000 years ago yet another population Branch out from our own, heading West and South into greater Africa. It is likely that this subspecies of Heidelberg assimilated the aforementioned African archaic that had been living there since 1.3 million years ago, as introgression from the group in pygmies often contains these older genes as well.
600,000 years ago another group split off from Heidelberg, heading East. Their genes are now only found at low levels in the Andaman islanders. The appearance of "pygmy heidelbergs" in northern India around 150,000 years ago may herald their arrival.
400000 years ago one group of Heidelberg left our gene pool and migrated into Subsaharan or West Africa and is represented by the later introgressed Y Haplogroup A00, which is today only found in a handful of people with ancestry from that region. It is separated by 200 thousand years from the haplogroups of the rest of modern man, suggesting 200 thousand years of haplogroup extinctions from which only two haplogroups among millions survived. Shortly after the divergence of this haplogroup, the tallest examples of Classic Heidelberg appear in Africa in the form of Rhodesian Man and Homo Sapiens Idaltu. Some populations are thought to have averaged 7 foot among males. African Ergaster happens to be the tallest member of the Genus Homo before African Heidelbergs such as Rhodesian man and Idaltu (all are 6'1+), while non-African Heidelbergs are the same height as the erectus that went before them (Antecessor and Los huesos are 6', all other non-African Erectus and "Heidelberg" range between 3' and 5'10, with 5'10 being an extreme rather than a population average). In evolutionary thought, it is considered easier to shrink than to grow, although I would guess that environment, gene exchange, and circumstance might also have something to do with it.
The last divergence from our from own direct ancestral population (that we know about) occurred between 250 and 350 thousand years ago and is evidenced in the mtDNA of an anatomically modern human dated 40-60 thousand years ago in Australia known as Mungo Man.
Anatomically humans show up in the fossil record until 160,000 years ago, and they are all in Africa, which might seem to contradict the DNA. Research agree that though they were both homo Sapien sapiens, neither our Mito eve nor that of Mungo Man were likely anatomically modern. Some hold that anatomically modern Homo Sapien Sapiens don't actually appear until around 120,000 years ago or later. How can it be that two seperate gene pools that were not anatomically modern to begin with both become anatomically modern by 45000 years ago? For that matter, since Y Haplogroup A00 split off from our own 400,000 years ago, when Homo Sapien Sapiens wasn't even around, how can A00 people living today be anatomically modern?
The answer is interbreding between the groups. According to john Hawks and the bBC program first Peoples, Anatomically modern humans formed from genetic interchange between several different groups of heidelbergensis. We know this because different groups of Archaic Humans living in or near Africa from 350 thousand to 90 thousand years ago have different pieces of the puzzle that coalesced to creat anatomically modern human morphology. The group that included the artist formerly known as Y DNA Adam, the group that contained Y Haplogroup A00, the group that contained Mitochondrial Eve, and the group that contained LM3 eve were all subgroups of Heidelberg, with different traits, and they came together to form the traits inherent in every living person today. Y Adam and Mito Eve just won the lineage marker part of the genome, there is no reason to believe that other groups didn't win other parts. Any introgression from the LM3 or A00 populations that happened before anatomically modern human came to be would not be seen as introgression, because everybody on Earth has them today. Too, unlike introgression from neanderthals and Denisovans, nuclear material from the A00 and LM3 populations would only be separated from our own nuclear material by a couple of hundred thousand years or less, and equidistant from that of Neanderthals and Denisovans, so there is even the possibility of introgression after the time in which they coelesced into AMH. We even have evidence for that in the form of a erroneous insert into 39% of modern human's own chromosome 11 that looks just like mtDNA from LM3's Eve. If not for this insert, scientists would not know that our own ancestors even bred with our Pacific sister species, leaving the possibility that there were still other extinct populations of Homo sapien sapiens and AMH that bred with us but remain undetected.
It is even a possibility that all of our Archaic Introgression stems from these "Ghost Population" having assimilated neanderthals, denisovans, and others before we assimilated them in turn.
The extreme of this is that all members of the Genus Homo should be considered Homo sapien sapiens, but because there was likely limited fertility between some of these groups and because there is cause for separation on anatomical grounds, not to mention the unlikelyhood that a modern human could successfully mate with a cloned Homo Habilis, I tend to disagree with this preference. However the various species and subspecies of the genus homo world without a doubt a part of an extended gene pool promise augmented and recombined many times over the past 3.5 million years.
90,000 years ago, LM3 may even still been a part of our own gene pool, and A00's population may not have yet recombined with it yet.
That's when a cold spell started pushing neanderthals into the Middle East and split our population into at least two parts. LM3 and possibly several Y and mtDNA haplogroups that survived into the present (Y Hap C and mtDNA M?) survived in South China where there is presence is known from 120,000 years ago, while the descendents of mito Eve (us) were limited to Africa. In a territory from Africa to Pakistan, and maybe into India, Neanderthal ruled and assimilated the AMH and Heidelberg remained in the region for the next 50 thousand years. we have evidence of this in not only the fossil record but from the introgressed genes of homo sapien sapiens found in the fossils of this self-same group of neanderthals, and their introgression into us.
This same cold spell may have opened up the land bridge to the Andaman Islands for the Early Heidelberg that is Andamanese Archaic Introgression. This subgroup of Heidelberg branched off from our own ancestors 600,000 years ago. By 150,000 years ago, a "pygmy" form of Heidelberg shows up in Northern India and lives for the next 30 thousand years or more alongside a taller but more gracile subgroup that had lived there for at least 600,000 years before. Their size and certain anatomical traits are almost identical to Andaman Islanders. Andaman islander Introgression may even be a combination of these two people's genes, with the 600,000 year old fossils, tools and introgressed genes being the taller Heidelbergs and the 150,000 year old tools and fossils being the Neanderthal Introgression in Andaman Islanders. It may also be that this group included women of the LM3 haplogroup, or a ghost population of Hss that carried the LM3 insert, because Andaman Islanders have it too. It could even have included Y Haplogroup C and D and mtDNA M, but this is doubtful, and here is the reason: introgression is only a fraction of Andaman Islander DNA. The vast majority of it is from the Homo Sapien Sapiens genome, and their maternal and paternal haplogroups are from within the even more finite branches of Y haplogroup A and mtDNA Eve. More importantly, they have all of the "updates" that make us human, unlike Indian Pygmy Heidelberg/Neanderthal or any Hominid that branched off from us 600,000 years ago would have. The dates just don't compute- C,D, and M are only around 60 thousand years old. their age would have to be multiplied by three to comply with the dates of the first pygmy Heidelberg's arrival in Northern India! Furthermore, the Andaman islanders are thought to have obtained their 600,000 year old genes only 35,000 years ago, and their Neanderthal genes not long before....and certainly not over 125,000 years before.
But what if the original population from which Andaman Islanders received their introgression was part Hss in the first place? What if in addition to neanderthal and 600,000 year old "Heidelberg genes" it also contained an equal or greater portion of genes from LM3 or Mito Eve? What if it contained Y Haplogroup A0? What if the shared cultural and physical traits of South East Asian Negritos and African pigmy's, such as steatopygia and the similarities between the two groups in cranial morphology as reported in the last study on the subject conducted in 1973.
What if the biggest difference between the negritos and pygmies and ourselves isn't the amount of neanderthal Introgression or admixture from 600,000 and 700,000 year old early heidelbergs. Maybe it's that they were never fully assimilated by the taller group of Homo Sapien sapiens who that once shared an enviroment with in the original homeland of Homo Sapien Sapiens, two populations enjoying two different niches and occasionally exchanging genes, until the Neanderthals drove both of us out.
It may even be that Y haps C and D and mtDNA M in Andaman Islanders comes from a handful of shipwrecked sailors who entered the gene pool sometime between 35,000 and 10,000 years ago.
More likely, however, is that they a group of Heidelberg/Neanderthal/Hss pygmies lived in Northern India, Southern China, and Indochina from 150,000 years ago to at least 45,000 years ago and shared the landscape and sometimes exchanged genes with a taller hybrid of heidelberg/ Hss hybrid that had experienced less introgression from Neanderthals. I say this because the taller specimens linked to Hss from this time and era are more gracile, and because the probable descendant Mungo Man is hyper-gracile and has supposedly less neanderthal introgression than other Hss specimens outside of Africa from a similar date.
An interesting sidenote is that Hss from both Japan and Australia during this period exhibit the practice of tooth knocking. The Japanese Hss have large denisovan like teeth and are Hyper-Robust like Denisovans and they are short and robust like the Indian Heidelberg pygmies, but the Australians are tall and lean. I suspect that the practice may have resulted from a problem with tooth development during adolescence in hybrids of the two groups, or in hybrids of the two groups and neanderthals or Asian hominids. The practice continued in Japan until the Neolithic Jomon Period, in Australia into the present, and back-migrated into Africa about 20,000 years ago within the late Aterian or Ibero-Maurusian cultures, possibly with Y Haplogroup DE and MtDNA M.
So it looks like we had two groups of Homo sapiens sapiens living in an area africa to indochina from around 150,000 years ago to 90,000 years ago. One exploded plains, savannas, and grasslands and was tall and gracile. The other exploited forests and was short and stocky. Ice sheets and Neanderthals kept them out of Europe and Central Asia and something kept them out of most of India, likely something closely related to neanderthals judging my the Mousterian tools found there. Something was also keeping both groups of Hss out of Sub-Saharan and West Africa, where A00 and Archaic African Introgression were waiting to be assimilated by people living there today. Indeed, archaic fossils persists in west Africa until 10,000 years ago.
A cold spell drove the Neanderthals into SW Asia and the Middle East, and separated the genome of LM3 and Negritos from the genome of Mito Eve and Pygmies. Before 90000 years ago, a few fossils of Homo Sapien sapiens in East Africa, China, and the Levant exhibit chins. After 90000 years ago, chins begin to pop up all over Africa and in Southern Neanderthals too. Some African examples dated between 70 thousand and 90 thousand years ago were considered neanderthals until the 1980s, when they were reassigned the Homo Sapien Sapiens classification due to a number of modern features. It looks for all the world like Neandertha/Hss hybrids from the borders of Africa and SW Asia are beginning an assimilation of the entire continent. The dates match up much well with the back-migration of Y Haplogroup BT, and with a second wave of DE and/or its offspring, basal y haplogroup E. The only Africans who seem to have escaped this assimilation are the pygmies, who likely had higher Malaria immunity due to their archaic introgression, and those who mixed with a population of Heidelbergs that contained A00. Both happened in Subsaharan or West Africa during the Upper Pleolithic, probably around 37,000 years ago.
Today, back-migrated Y haplogroup E makes up over 70% of men native to Africa and the vast majority of Africans outside of Subsaharan Africa possess neanderthal introgression. It is mainly only pygmies who have managed to avoid it, and they have introgression that likely serves them far better than a cold-bred hominid's would.
From 70000 years ago to about 45,000 years ago the Y Hap DE neanderthal/Hss hybrid males dominated most of SW Asia, Northern and Eastern Africa, and the Middle East. Their original mtDNA counterpart was likely N, but they also picked up some mtDNA M, possible from the North Indian Pygmies. Sometime around 20,000 years ago a group of them brought tooth-knocking and mtDNA haplogroup M into Africa, and at some point around this same time one of their groups reached the Andaman Islands carrying the assimilated genes of proto-negritos.
This may have been the result of the population in which de was dominant being split in half, as the genome of homo sapiens sapiens had been split in half by neanderthals before. The reason for this was either the emptying of Central asia and the middle East due to climactic conditions or the incursion of another hybrid group from India, the one that contained Y haplogroup FC.
The dispersal of Y Haplogroup C, probably from South Asia, begins around 60,000 years ago and continues into Eurasia and South east Asia. The dispersal of F begins about 45,000 years ago and does the same. My best guess is that C represents those South Asian Hss that bred with the short woodland Neanderthals of South Asia and that F represents those who bred with the taller neanderthals of South Asia. Hybrid vigor allowed both groups' expansions into larger India, assimilating the early neanderthals there who had already mixed with an incursion Microcephalin D hominids that were expanding in the wake of lower sea levels from South east Asia. This is why the Microcephalin D gene is most prevalent in populations dominated by F haplogroups and mostly absent in African and Asian groups dominated by E and D, the offspring of Y haplogroup DE. Assimilation of the 3 million year divergent Homo Varana probably didn't happen until 21 thousand years ago at the last glacial maximum, when Adam's bridge to Sri Lanka became dry again. The resulting hybrids are known as Balangoda man, and some of them seemed to have moved to SW of India itself where they live to this day.
The push of F and C into central Asia pushed neanderthal East and West, and in the the East they began assimilating their sister species Denisovan, who had already begun assimilating Early Homo some 300 thousand years before. Neanderthal like skulls have been found in Mongolia and near Biejing with neanderthal and other archaic features with dates as late as 9000 B.C. The Red Deer Cave people possess triats from Erectus, neanderthal, and Hss...along with a few that only they can boast. Neanderthal stone balls, previously only found in Europe, show up around 40 thousand years ago in China and continue into South East Asia no later than 27,000 years ago. This is why Northern Chinese, native Amnericans. and Papuans have more Neanderthal DNA than most Europeans: Neanderthal assimilated most of the Denisovan and the Microcephalin D populations before we got there, along with much of the genome of our Mungo Man brother LM3.
It also seems that neanderthals and other hominids liked our females better than their own, as their is evidence for Hss introgression into Neanderthals in some Asian specimens and our mtDNA haplogroups seem to preceed us into certain areas, such as Tibet and Europe, long before any of our surviving Y haplogroups get there. When we took the world back from Neanderthal, we must not have killed the women, or at least we didn't kill the women who looked like us.
But why would all of these dominant neanderthal hybrids who took over the world except for african pygmies always be from an Hss haplogroup rather than a neanderthal one? It may have something to do with mammal pattern infertility. In mammalian hybrids, the females are more likely to be fertile than the males. The lack of neanderthal genital introgression makes it probable that hybrids who inherited the blackneanderthal reproductive genes rather than their Hss parent's were less successful in producing offspring. The other genes missing from the Neanderthal genome in modern introgression have to do with eyesight and may have been a factor in the size of the occipital bun, which could result in birth complications. if Neanderthals were Blood Type O Negative as is supposed, it may be that they could only have one child with an Hss wiothout the aid of modern medicine. It might also be because homo Sapien sapiens was the first hominid that did not need to go into heat in order to procreate. This could explain other subgroups choosing Hss women over their own and the rapid expansion of Hss genes after the Toba catastrophe. We simply outbred them. It could also be that Hss has a more neonatal look than any hominid that preceded it. We were originally pygmies that were just too cute to kill, and our women were smoking hot!
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